 | Liu, Y.H., T. Kuwada, and Y. Koya, 2008, Gonadal sex differentiation in Ayu, Plecoglossus altivelis altivelis., Japanese Journal of Ichthyology. |
| Masuda, H., K. Amaoka, C. Araga, T. Uyeno, and T. Yoshino, 1984, The Fishes of the Japanese Archipelago ., Tokai Univ. Press. Fish. Japanese Arch. |
| 沈世傑 編 Shih-Chieh Shen ed., 1993, 臺灣魚類誌 Fishes of Taiwan, 國立臺灣大學動物學系印行 |
| Nakabo T., 2002, Fishes of Japan - with pictorial keys to the species, English edition., Tokai University. |
| Nakabo T., 2002, Fishes of Japan- with pictorial keys to the sprcies, English edition., Tokai University. |
